Sherwood Forest coleoptera - species gains and losses
In what could quite possibly be a case of 'out with the old and in with the new', there's been some striking changes to Sherwood Forest's beetle fauna over the past 100 years.

This page lists both those beetles unrecorded within the area known as the Sherwood Forest National Nature Reserve since 1921 and those recorded new to the area since 2000. It's quite some turnover really and it would be interesting to know if such a list of recently added species is actually quite typical of other UK sites.

Sherwood is fortunate in that it's invertebrate fauna over most orders, is generally well recorded, documented and accessible. This makes things easier for those of us interested in the variances brought about over the course of a century, through changes to climate and an endless range of other possible influencing factors affecting many species either positively or negatively.

The effects for inducing the sudden or gradual decline, or expansion of a particular species, are probably several in the vast majority of cases and the cause of decline or loss, can be beyond our understanding of that species.

Just having the correct habitat is often not enough to rescue a species from catastrophic decline and a number of beetles have gone that way over the years.
Sherwood Forest NNR species list total and changes to the species list

As of December 31st 2020, the Sherwood Forest National Nature Reserve coleoptera list stood at an approximate total of 1,201 species. 20 years ago (at the start of this century) the species list totalled 922, which was 84 species up on the 824 total reached at the end of 1921, some five years after J.W. Carr published his 1916 book 'The invertebrate Fauna of Nottinghamshire'. How lucky Nottinghamshire is to have such a book for the researchers among us.

The Sherwood Forest list should be considered healthy for an area of it's size, especially as it lacks areas of deciduous woodland with a more diverse range of trees and shrubs and a large wetland area with good marginal habitat. Species-wise, it has seen considerable changes since the earliest documented records came in the late 1800's and of those species recorded up to 1921, approximately 297 have not been recorded since. Although beetles have been regularly 'rediscovered' at Sherwood after having gone unrecorded for a century, it is highly likely that many have indeed been lost from the area over time. On the flip side, a total of 293 previously unrecorded species, have been gained since 2000 alone. This rather high total, is the result of Sherwood's most intense period of coverage and recording effort geared towards coleoptera.  
  The total includes 14 aquatic species recorded new to the Sherwood Forest NNR post-2000, from the families Haliplidae, Noteridae, Hygrobidae and Disticidae and many more had been added during the latter part of the 1990's. For some reason, no records of these or any other aquatic coleoptera were recorded (or presented) from anywhere in the Sherwood Forest area in Carr's 1916 book. Why this is remains unknown and something of a mystery really, as there were obviously some water bodies present over the area of the NNR a century or more ago.

Surveys, recorders and species additions

The credit of Sherwood Forest even having a list of aquatic coleoptera, is really down to the enormous recording efforts of Bob Merritt. Bob singly handedly mapped Nottinghamshire's aquatic coleoptera during the late 1990's and early part of the 2000's and very kindly gave Dilys and myself a spreadsheet containing his records. Bob wanted his records 'used' and of course we gratefully obliged. An example of the results of Bob Merritt's recording is shown in the map for Ilybius fuliginosus (Fabricius, 1792) on the left. All bar about a dozen of the grid squares are attributable to Bob's recording. Quite a remarkable achievement.

Bob Merritt left Nottinghamshire after eventually adding some 63 (mostly aquatic) species new to the NNR. Dilys and myself contributed 102 new species to the NNR list during many visits between 2008 and 2015, but many other new species (with totals in brackets) have been added following surveys conducted on behalf of Natural England East Midlands by M.L. Denton, E.J. Green, D.G. Hemingway, R.J. Marsh and D. Sharp, D. in 2002 (55sp) and more recently by a Natural England Survey Team of J. Hackman, J. Webb, A. Drewitt, J. Small and N. Button et al in 2018 (12sp).

Earlier site surveys included Derek Lott's 1998 survey (23sp), followed quickly by Tony Drane's 2000 survey (40sp), Keith Alexander's 2009/2010 survey (5sp) and Paul and Helen Brock and John and Denise Bingham's site visit/survey in 2018 (2sp)

General invertebrate surveys of Sherwood Heath by A. Godfrey and B. Marsh, commisioned by the Sherwood Forest Trust in the early 2000's, were a source of 36 new species for the Sherwood Forest NNR. Importantly, these surveys added many non-saproxylic species and most of the species found new this century, have largely been of this nature. This has placed a greater emphasis on Sherwood Forest holding a wider range of coleoptera, rather than just being considered important for it's saproxylic species. Sherwood is much more than that.
It's also important to remember the enormous contributions made by those often forgotten recorders, who covered Sherwood Forest throughout the latter decades of the 20th century. Colin Johnson found 24 species new to the area over a number of years between 1960 and 1989, while Peter Skidmore recorded another nine new species and F.A. Hunter six. Allan and Annette Binding added a further ten species during visits and surveys crossing both the previous and current centuries, while Adrian Dutton (a regular recorder of the NNR's coleoptera for a number of years) has added at least eight new species and a number of 'refounds' during his visits.

Species for which there have been no Sherwood Forest NNR records since 1921

The following list of beetles have not been recorded from anywhere within the Sherwood Forest NNR over the last 100 years. While many of those listed are small species, often from difficult to identify families or families out of favour with coleopterists today, it is always possible that they could be recorded at some time in the future with persistant recording effort. It has to be said however, that a great many of the species on this list are probably best considered lost and not being present at Sherwood Forest today. There are some species on the list though, which have declined nationally in recent decades and it may be that over time, some of these will return when conditions are more suited.
  The cause of a species being lost from a site can be down to any number of causes and influencing factors; either through natural means such as microclimatic or global climate changes, or the loss of habitat through human activity such as commercial forestry, an over-zealous scrub/tree removal program, or just inconsiderate and poor site management. 

Contrary to popular general opinion, there is plenty of standing dying, or dead wood habitat at Sherwood Forest Country Park. But if trees keep being removed as part of a thinning program aimed at increasing veteran tree longevity and to ensure 'healthy' woodland, there is every chance that this will potentially lead to a lack of standing dying/dead wood habitat. This is being compensated for by the abhorrent practice of 'veteranising' trees to speed up the aging process. It's just arboreal euthanasia and it's needless.

To ensure a continuous supply of dying/deadwood habitat at Sherwood Forest, it would perhaps be better practice to remedy Sherwood's age-old problem of being a forest of trees with a limited age range. As a mature forest with a quite unique population of veteran trees, unless young Oak and the forgotten Silver Birch are allowed to grow now, there will be even less tree succession in 100/200 years from now and aren't we supposed to be protecting the forest for future generations?    
Ptiliidae has been one of the worst affected families in terms of species presumed as lost from the NNR. Some ten species have not been seen in 100 years, including four species of Acrotrichis. These seem to have been replaced by four new species of Acrotrichis since the turn of this century however and there must be a number of others awaiting discovery for anyone fancying something of a challenge.

Leiodidae is another family to have suffered a number of losses in Sherwood Forest, with 19 species not recorded since before 1921 and only five new species found since 2000. Despite members of this family being distinctive to assign to family level easily enough, identification to species level is not always easy and as they are often being found on or near carrion or fungi, perhaps aren't many coleopterist's first choice of study?

A total of 95 species from the extensive Staphylinidae family, have not been recorded since 1921, but Staphylinidae is another of those families often shunned by modern recorders, largely through the difficulties in species identification and the huge number of species the family contains. Other families and their corresponding number of lost species not seen since 1921, include Elateridae (5sp), Cantharidae (4sp), Nitidulidae (10sp), Monotomidae (6sp), Latridiidae (7sp), Cerambycidae (4sp), Chrysomelidae (23sp), Apionidae (14sp) and Curculionidae (30sp).
Carabidae Latreille, 1802
Bembidion femoratum Sturm, 1825
Poecilus kugelanni (Panzer, 1797)
Pterostichus diligens (Sturm, 1824)
Synuchus vivalis (Illiger, 1798)
Calathus erratus (Sahlberg, C.R., 1827)
Olisthopus rotundatus (Paykull, 1790)
Amara fulva (Müller, O.F., 1776)
Harpalus rubripes (Duftschmid, 1812)
Harpalus tardus (Panzer, 1796)
Bradycellus caucasicus (Chaudoir, 1846)
Anthracus consputus (Duftschmid, 1812)
Licinus depressus (Paykull, 1790)
Helophoridae Leach, 1815
Helophorus nubilus Fabricius, 1777
Helophorus rufipes (Bosc d'Antic, 1791)
Hydrophilidae Latreille, 1802
Cercyon analis (Paykull, 1798)
Cryptopleurum minutum (Fabricius, 1775)
Histeridae Gyllenhal, 1808
Abraeus granulum Erichson, 1839
Acritus nigricornis (Hoffmann, J., 1803)
Hypocaccus rugifrons (Paykull, 1798)
Carcinops pumilio (Erichson, 1834)
Margarinotus carbonaris (Hoffmann, 1803)
Margarinotus merdarius (Hoffmann, J., 1803)
Margarinotus purpurascens (Herbst, 1792)
Ptiliidae Erichson, 1845   
Ptenidium gressneri Erichson, 1845
Ptenidium turgidum Thomson, C.G., 1855
Euryptilium saxonicum (Gillmeister, 1845)
Ptiliolum marginatum (Aubé, 1850)
Ptinella denticollis (Fairmaire, 1858)
Ptinella limbata (Heer, 1841)
Acrotrichis atomaria (De Geer, 1774)
Acrotrichis brevipennis (Erichson, 1845)
Acrotrichis montandonii (Allibert, 1844)
Acrotrichis thoracica (Waltl, 1838)
Leiodidae Fleming, 1821  
Leiodes calcarata (Erichson, 1845)
Leiodes dubia (Kugelann, 1794)
Leiodes obesa (Schmidt, W.L.E., 1841)
Leiodes oblonga (Erichson, 1845)
Colenis immunda (Sturm, 1807)
Agathidium convexum Sharp, 1866
Agathidium marginatum Sturm, 1807
Agathidium rotundatum (Gyllenhal, 1827)
Nargus wilkinii (Spence, 1813)
Choleva angustata (Fabricius, 1781)
Sciodrepoides fumatus (Spence, 1813)
Catops chrysomeloides (Panzer, 1798)
Catops coracinus Kellner, 1846
Catops grandicollis Erichson, 1837
Catops kirbii (Spence, 1813)
Catops morio (Fabricius, 1787)
Ptomaphagus medius Rey, 1889
Ptomaphagus varicornis (Rosenhauer, 1847)
Leptinus testaceus Müller, P.W.J., 1817
Staphylinidae Latreille, 1802 
Eusphalerum luteum (Marsham, 1802)
Eusphalerum torquatum (Marsham, 1802)
Omalium allardi Fairmaire & Brisout de Barneville, 1859
Omalium excavatum Stephens, 1834
Omalium exiguum Gyllenhal, 1810
Omalium italicum Bernhauer, 1902
Omalium oxyacanthae Gravenhorst, 1806
Phloeonomus pusillus (Gravenhorst, 1806)
Phyllodrepa salicis (Gyllenhal, 1810)
Megarthrus bellevoyei (Saulcy, 1862)
Megarthrus denticollis (Beck, 1817)
Proteinus crenulatus Pandellé, 1867
Arrhenopeplus tesserula Curtis, 1828
Euplectus sanguineus Denny, 1825
Euplectus signatus (Reichenbach, 1816)
Bryaxis curtisi (Leach, 1817)
Tychus niger (Paykull, 1800)
Lordithon exoletus (Erichson, 1839)
Mycetoporus angularis Mulsant & Rey, 1853
Mycetoporus longulus Mannerheim, 1830
Mycetoporus rufescens (Stephens, 1832)
Cilea silphoides (Linnaeus, 1767)
Tachinus proximus Kraatz, 1855
Tachyporus pusillus Gravenhorst, 1806
Trichophya pilicornis (Gyllenhal, 1810)
Aleochara lanuginosa Gravenhorst, 1802
Aleochara lata Gravenhorst, 1802
Aleochara moerens Gyllenhal, 1827
Aleochara moesta Gravenhorst, 1802
Tinotus morion (Gravenhorst, 1802)
Atheta boletophila (Thomson, C.G., 1856)
Atheta divisa (Märkel, 1844)
Atheta euryptera (Stephens, 1832)
Atheta fungicola (Thomson, C.G., 1852)
Atheta liturata (Stephens, 1832)
Atheta ravilla (Erichson, 1839)
Badura macrocera (Thomson, C.G., 1856)
Dimetrota atramentaria (Gyllenhal, 1810)
Liogluta longiuscula Thomson, C.G., 1858
Microdota atricolor (Sharp, 1869)
Microdota benickiella (Brundin, 1948)
Microdota subtilis (Scriba, 1866)
Nehemitropia lividipennis (Mannerheim, 1830)
Pachyatheta cribrata (Kraatz, 1856)
Traumoecia picipes (Thomson, C.G., 1856)
Autalia rivularis (Gravenhorst, 1802)
Cordalia obscura (Gravenhorst, 1802)
Gyrophaena affinis Mannerheim, 1830
Gyrophaena fasciata (Marsham, 1802)
Gyrophaena gentilis Erichson, 1839
Gyrophaena nana (Paykull, 1800)
Oligota inflata (Mannerheim, 1830)
Oligota picipes (Stephens, 1832)
Oligota pusillima (Gravenhorst, 1806)
Pella limbata (Paykull, 1789)
Myllaena fowleri Matthews, A., 1883
Haploglossa villosula (Stephens, 1832)
Ilyobates nigricollis (Paykull, 1800)
Ischnoglossa prolixa (Gravenhorst, 1802)
Placusa tachyporoides (Waltl, 1838)
Anotylus inustus (Gravenhorst, 1806)
Oxytelus fulvipes Erichson, 1839
Oxytelus sculptus Gravenhorst, 1806
Platystethus arenarius (Fourcroy, 1785)
Eutheia linearis Mulsant, 1861
Neuraphes angulatus (Müller, P.W.J. & Kunze, 1822)
Scydmoraphes sparshalli (Denny, 1825)
Stenichnus godarti (Latreille, 1806)
Stenus nanus Stephens, 1833
Lathrobium elongatum (Linnaeus, 1767)
Othius angustus Stephens, 1833
Othius laeviusculus Stephens, 1833
Bisnius cephalotes (Gravenhorst, 1802)
Bisnius puella (Nordmann, 1837)
Gabrius trossulus (Nordmann, 1837)
Philonthus addendus Sharp, 1867
Philonthus albipes (Gravenhorst, 1802)
Philonthus cruentatus (Gmelin in Linnaeus, 1790)
Philonthus marginatus (Müller, O.F., 1764)
Philonthus politus (Linnaeus, 1758)
Philonthus ventrialis (Gravenhorst, 1802)
Heterothops dissimilis (Gravenhorst, 1802)
Quedius boops (Gravenhorst, 1802)
Quedius cinctus (Paykull, 1790)
Quedius fulgidus (Fabricius, 1792)
Quedius levicollis (Brullé, 1832)
Quedius picipes (Mannerheim, 1830)
Quedius puncticollis (Thomson, C.G., 1867)
Quedius semiaeneus (Stephens, 1833)
Quedius semiobscurus (Marsham, 1802)
Quedius truncicola Fairmaire & Laboulbène, 1856
Ocypus opthalmicus (Scopoli, 1763)
Staphylinus caesareus Cederhjelm, 1798
Megalinus glabratus (Gravenhorst, 1802)
Xantholinus elegans (Olivier, 1795)
Geotrupidae Latreille, 1802
Trypocopris vernalis (Linnaeus, 1758)
Scarabaeidae Latreille, 1766 
Aphodius coenosus (Panzer, 1798)
Scirtidae Fleming, 1821 
Elodes minuta (Linnaeus, 1767)
Cyphon coarctitus Paykull, 1799
Cyphon variabilis (Thunberg, 1787)
Prionocyphon serricornis (Müller, P.W.J., 1821)
Byrrhidae Latreille, 1804 
Byrrhus pustulatus (Forster, 1770)
Dryopidae Billberg, 1820 (1817) 
Dryops ernesti des Gozis, 1886
Elateridae Leach, 1815
Calambus bipustulatus (Linnaeus, 1767)
Aplotarus incanus (Gyllenhal, 1827)
Athous campyloides Newman, 1833
Adrastus rachifer (Fourcroy, 1785)
Agriotes lineatus (Linnaeus, 1767)
Ampedus cinnabarinus (Eschscholtz, 1829)
Cantharidae Imhoff, 1856 (1815) 
Podabrus alpinus (Paykull, 1798)
Cantharis paludosa Fallén, 1807
Malthodes fibulatus Kiesenwetter, 1852
Malthodes pumilus (Brébisson, 1835)
Bostrichidae Latreille, 1802
Lyctus brunneus (Stephens, 1830)
Ptinidae Latreille, 1802
Ptinus subpilosus Sturm, 1837
Stegobium paniceum (Linnaeus, 1758)
Anitys rubens (Hoffmann, J., 1803)
Cleridae Latreille, 1802 
Necrobia ruficollis (Fabricius, 1775)
Necrobia rufipes (De Geer, 1775)
Necrobia violacea (Linnaeus, 1758)
Dasytidae Laporte, 1840
Dasytes plumbeus (Müller, O.F., 1776)
Kateretidae Kirby, 1837
Kateretes pusillus (Thunberg, 1794)
Brachypterus urticae Nettle Pollen Beetle (Fabricius, 1792)
Nitidulidae Latreille, 1802 
Carpophilus mutilatus Erichson, 1843
Epuraea guttata (Olivier, 1811)
Epuraea longula Erichson, 1845
Epuraea silacea (Herbst, 1783)
Meligethes difficilis (Heer, 1841)
Meligethes morosus Erichson, 1845
Meligethes ochropus Sturm, 1845
Meligethes viridescens (Fabricius, 1787)
Nitidula bipunctata (Linnaeus, 1758)
Omosita depressa (Linnaeus, 1758)
Monotomidae Laporte, 1840 
Rhizophagus oblongicollis Blatch & Horner, 1892
Rhizophagus parallelocollis Gyllenhal, 1827
Rhizophagus picipes (Olivier, 1790)
Monotoma conicicollis Guérin-Méneville, 1837
Monotoma longicollis (Gyllenhal, 1827)
Monotoma testacea Motschulsky, 1845
Silvanidae Kirby, 1837
Oryzaephilus surinamensis Saw-toothed Grain Beetle (Linnaeus, 1758)
Cryptophagidae Kirby, 1837
Cryptophagus pubescens Sturm, 1845
Cryptophagus setulosus Sturm, 1845
Antherophagus similis Curtis, 1835
Atomaria apicalis Erichson, 1846
Atomaria nigriventris Stephens, 1830
Erotylidae Latreille, 1802 
Dacne rufifrons (Fabricius, 1775)
Bothrideridae Erichson, 1845 
Oxylaemus cylindricus (Creutzer in Panzer, 1796)
Coccinellidae Latreille, 1807  
Scymnus auritus Thunberg, 1795
Scymnus nigrinus Kugelann, 1794
Platynaspis luteorubra (Goeze, 1777)
Coccinella undecimpunctata 11-spot Ladybird Linnaeus, 1758
Corylophidae LeConte, 1852
Orthoperus atomarius (Heer, 1841)
Orthoperus atomus (Gyllenhal, 1808)
Orthoperus brunnipes (Gyllenhal, 1808)
Orthoperus corticalis (Redtenbacher, 1849)
Latridiidae Erichson, 1842
Latridius porcatus Herbst, 1793
Enicmus brevicornis (Mannerheim, 1844)
Deinerella filiformis (Gyllenhal, 1827)
Corticaria crenulata (Gyllenhal, 1827)
Corticaria fulva (Comolli, 1837)
Corticaria dubia Dajoz, 1970
Corticaria punctulata Marsham, 1802
Mycetophagidae Leach, 1815  
Mycetophagus piceus (Fabricius, 1777)
Mycetophagus quadripustulatus (Linnaeus, 1761)
Ciidae Leach, 1819
Ennearthron comutum (Gyllenhal, 1827)
Tenebrionidae Latreille, 1802  
Corticeus linearis (Fabricius, 1790)
Diaperis boleti (Linnaeus, 1758)
Oedemeridae Latreille, 1810 
Ischnomera sanguinicollis (Fabricius, 1787)
Scraptiidae Gistel, 1848  
Scraptia fuscula Müller, P.W.J., 1821
Anaspis lurida Stephens, 1832
Cerambycidae Latreille, 1802 
Stictoleptura scutellata (Fabricius, 1781)
Alosterna tabacicolor (De Geer, 1775)
Gracilia minuta (Fabricius, 1781)
Stenostola dubia (Laicharting, 1784)
Chrysomelidae Latreille, 1802
Bruchidius villosus (Fabricius, 1792)
Cryptocephalus nitidulus Fabricius, 1787
Chrysolina sturmi (Westhoff, 1882)
Gastrophysa polygoni (Linnaeus, 1758)
Hydrothassa marginella (Linnaeus, 1758)
Prasocuris junci (Brahm, 1790)
Phratora vitellinae (Linnaeus, 1758)
Luperus flavipes (Linnaeus, 1767)
Luperus longicornis (Fabricius, 1781)
Phyllotreta vittula (Redtenbacher, 1849)
Longitarsus atricillus (Linnaeus, 1761)
Longitarsus exoletus (Linnaeus, 1758)
Longitarsus ganglbaueri Heikertinger, 1912
Longitarsus gracillis Kutschera, 1864
Longitarsus pratensis (Panzer, 1794)
Altica helianthemi (Allard, 1859)
Neocrepidodera ferruginea (Scopoli, 1763)
Derocrepis rufipes (Linnaeus, 1758)
Mantura rustica (Linnaeus, 1767)
Apteropeda orbiculata (Marsham, 1802)
Psylliodes cuprea (Koch, J.D.W., 1803)
Psylliodes luteola (Müller, O.F., 1776)
Cassida viridis Linnaeus, 1758
Nemonychidae Bedel, 1882
Cimberis attelaboides (Fabricius, 1787)
Anthribidae Billberg, 1820 
Anthribus nebulosus (Forster, 1771)
Rhynchitidae Gistel, 1848
Involvulus cupreus (Linnaeus, 1758)
Neocoenorrhinus germanicus (Herbst, 1797)
Neocoenorrhinus pauxillus (Germar, 1824)
Attelabidae Billberg, 1820 
Apoderus coryli (Linnaeus, 1758)
Apionidae Schönherr, 1823 
Apion cruentatum Walton, J., 1844
Aspidapion aeneum (Fabricius, 1775)
Aspidapion radiolus (Marsham, 1802)
Ceratapion carduorum (Kirby, W., 1808)
Eutrichapion ervi (Kirby, W., 1808)
Holotrichapion aethiops (Herbst, 1797)
Holotrichapion pisi (Fabricius, 1801)
Oxystoma craccae (Linnaeus, 1767)
Pirapion immune (Kirby, W., 1808)
Ischnopterapion loti (Kirby, 1808)
Ischnopterapion virens (Herbst, 1797)
Protopirapion atratulum (Germar, 1817)
Protapion assimile (Kirby, W., 1808)
Protapion difforme (Germar, 1818)
Erirhinidae Schönherr, 1825
Procas picipes (Marsham, 1802)
Curculionidae Latreille, 1802
Curculio betulae (Stephens, 1831)
Cleopus pulchellus (Herbst, 1795)
Dorytomus tortrix (Linnaeus, 1761)
Orchestes alni (Linnaeus, 1758)
Orchestes calceatus (Germar, 1821)
Orchestes testaceus (Müller, O.F., 1776)
Tachyerges salicis (Linnaeus, 1758)
Ceutorhynchus alliariae Brisout de Barneville, H., 1860
Ceutorhynchus assimilis (Paykull, 1792) sensu.stricto.
Ceutorhynchus obstrictus (Marsham, 1802)
Ceutorhynchus pyrrhorhynchus (Marsham, 1802)
Coliodes transversealbofasciatus (Goeze, 1777)
Datonychus arquatus (Herbst, 1795)
Phytobius leucogaster (Marsham, 1802)
Rhyncolus ater (Linnaeus, 1758)
Otiorhynchus ligneus (Olivier, 1807)
Phyllobius glaucus (Scopoli, 1763)
Phyllobius viridicollis (Laicharting, 1781)
Barypeithes araneiformis (Schrank, 1781)
Sitona hispidulus (Fabricius, 1777)
Sitona puncticollis Stephens, 1831
Hypera suspiciosa (Herbst, 1795)
Hypera venusta (Fabricius, 1781)
Magdalis armigera (Fourcroy, 1785)
Scolytus ratzeburgi Birch Bark Beetle Janson, E.W., 1856
Orthotomicus laricis (Fabricius, 1792)
Hylesinus varius Ash Bark Beetle (Fabricius, 1775)
Species recorded as new to the Sherwood Forest NNR since 2000

Of the following 293 species for which there appear to be no documented records from Sherwood Forest prior to 2000, many will come as a surprise to the experienced coleopterist who happens to read this. The list includes a number of beetles which in all honesty, would be regarded as standard 'list fillers' for any invertebrate survey today, and there seems to be no real reason why they haven't been recorded within the Sherwood Forest NNR until recently. Quite astonishing is the number of new Carabidae recorded new to the NNR, even though Carabidae is among the most popular families with coleopterists today.
A total of 39 new Carabids includes ten from the genus Amara alone and none of which are sufficiently rare enough to get really excited about. Why these weren't picked up before is strange, as none of them come immediately to mind as having undergone recent range expansions in the past 20 or so years.

Possibly it's because much of the recent coleoptera surveying, has been largely geared towards Sherwood's saproxylic species and that few individual recorders have devoted much time to recording here. Sherwood Forest is known ofr it's saproxylic invertebrates (not just some famous outlaw) and one of the reasons why it was originally awarded it's NNR status.

The 293 species does contain many new saproxylic beetles, including Agrilus sinuatus (Buprestidae) pictured right, Lymexylon navale (Lymexylidae), Orchasia minor and Melandrya caraboides (Melandryidae), Uleiota planata and Silvanus bidentatus (Silvanidae), Rhizophagus fenestralis (Monotomidae) and Cryptolestes duplicatus (Laemophloeidae). The latter beetle found in early 2020, is every bit the typical saproxylic beetle - dull, brown and small. However, excitingly it was the NNR's first recorded species from the family Laemophloeidae, as was Ochthebius minimus (Hydraenidae) which was another record attributable to Bob Merritt.
Four species of Acrotrichis (Ptiliidae) were added during the 2002 survey of Sherwood Forest Country Park and the Buck Gates area by M.L. Denton et al, but only one has been recorded again since. Acrotrichis and Ptiliidae are minute beetles generally (between 0.5 - 2mm in length) and although common in most leaf litter samples, are just too small for most people to bother with.

Another family not entirely popular with many coleopterists is Staphylinidae. This is a huge family with well over 1,000 UK species and probably more still  waiting to be discovered. There are many very small, similar species and the lack of a decent, readily available and up to date identification key to the whole family doesn't help. The recording of Staphylinids over the past century at Sherwood Forest is especially interesting to those who like their statistics, which is perhaps merely just the same as saying 'are interested'. The family's popularity at Sherwood, has certainly lessened over the past 100 years with 95 species either lost or unrecorded since 1921, compared to just 67 species gained.
  However, although the family's overall popularity has declined, a number of genera within the family are still popular with recorders and within the five genera which follow, Dilys and I certainly favoured Stenus, Philonthus and Quedius. Within these lost and gained Staphylinids are some interesting changes to at least five different genera, namely Tachyporus (1L/5G), Atheta (6L/3G), Stenus (1L/9G), Philonthus (6L/3G) and Quedius (9L/5G). Obviously 'L' means Lost and 'G' means Gained.

Observer preference or favouritism, certainly plays a part here and particularly with the genus Stenus. Seven out of the nine species gained since 2000, were once again attributable to the recording of Bob Merritt, but most of the other new species are from various commitioned surveys.

Compartmental grazing within the Sherwood Forest Country Park and on Budby South Forest by a combination of English Longhorn Cattle and Hebridean Sheep, has led to the largest increase in new species showed by Scarabaeidae. Prior to 2000, just six species were known compared to today's 24 and of the 24, personal favouritism towards the genus Aphodius has certainly been a factor in increasing the Sherwood species list.
Family favouritism also applies to Coccinellidae and it was surprising when originally compiling a species list for the NNR in 2008, that so many common and well recorded Ladybirds, hadn't actually been officially documented as having occurred at Sherwood, in the available media we had at the time. So to have species such as Tytthaspis sedecimpunctata 16-spot Ladybird, Psyllobora vigintiduopunctata 22-spot Ladybird and Adalia decempunctata 10-spot Ladybird as new to Sherwood post-2000, is rather ridiculous. The same might be thought about Halyzia sedecimguttata Orange Ladybird, but that's not so frequent over much of the NNR since most of the Sycamore's were removed.

But among the list of those species rather carelessly unrecorded, are some which never occurred anywhere near Nottinghamshire a number of years ago, but have moved north through the UK and appeared shortly after the turn of the present century. Harmonia axyridis Harlequin Ladybird is a classic example, as are the now ubiqutous Oedemera nobilis Swollen-thighed Beetle (one of two new Oedemeridae species) and Agelastica alni Alder Leaf Beetle (Chrysomelidae). But quite where Budby (and Newstead Abbey's) Meloe proscarabaeus Black Oil Beetle suddenly came from in 2011, still baffles to this day. Who knows just what will turn up next?
Haliplidae Aube 1836
Haliplus lineatocollis (Marsham, 1802)
Haliplus ruficollis (De Geer, 1774)
Noteridae Thomson, C.G., 1860
Noterus clavicornis (De Geer, 1774)
Hygrobiidae Erichson, 1837
Hygrobia hermanii Screech Beetle (Fabricius, 1775)
Dytiscidae Leach, 1815
Ilybius montanus (Stephens, 1828)
Ilybius quadriguttatus (Lacordaire, 1835)
Rhantus suturalis (MacLeay, 1825)
Liopterus haemorrhoidalis (Fabricius, 1787)
Disticus marginalis Great Diving Beetle Linnaeus, 1758
Hydroporus angustatus Sturm, 1835
Hydroporus melanarius Sturm, 1835
Suphrodytes dorsalis (Fabricius, 1787)
Hygrotus nigrilineatus (Steven, 1808)
Hyphydrus ovatus (Linnaeus, 1761)
Carabidae Latreille, 1802
Nebria salina Fairmaire & Laboulbène, 1854
Notiophilus substriatus Waterhouse, G.R., 1833
Trechus quadristriatus (Schrank, 1781)
Asaphidion curtum (Heyden, 1870)
Ocys harpaloides (Audinet-Serville, 1821)
Bembidion aeneum Germar, 1824
Bembidion articulatum (Panzer, 1795)
Bembidion guttula (Fabricius, 1792)
Bembidion obtusum Audinet-Serville, 1821
Poecilus cupreus (Linnaeus, 1758)
Poecilus versicolor (Sturm, 1824)
Pterostichus melanarius (Illiger, 1798)
Pterostichus niger (Schaller, 1783)
Pterostichus vernalis (Panzer, 1795)
Calathus fuscipes (Goeze, 1777)
Calathus melanocephalus (Linnaeus, 1758)
Laemostenus terricola (Herbst, 1784)
Anchomenus dorsalis (Pontoppidan, 1763)
Agonum gracile Sturm, 1824
Agonum muelleri (Herbst, 1784)
Amara communis (Panzer, 1797)
Amara convexior Stephens, 1828
Amara equestris (Duftschmid, 1812)
Amara eurynota (Panzer, 1796)
Amara familiaris (Duftschmid, 1812)
Amara lunicollis Schiødte, 1837
Amara ovata (Fabricius, 1792)
Amara plebeja (Gyllenhal, 1810)
Amara similata (Gyllenhal, 1810)
Amara tibialis (Paykull, 1798)
Harpalus affinis (Schrank, 1781)
Harpalus rufipalpis Sturm, 1818
Harpalus rufipes (De Geer, 1774)
Anisodactylus binotatus (Fabricius, 1787)
Bradycellus harpalinus (Audinet-Serville, 1821)
Bradycellus ruficollis (Stephens, 1828)
Acupalpus parvulus (Sturm, 1825)
Badister bullatus (Schrank, 1798)
Dromius angustus Brullé, 1834
Helophoridae Leach, 1815
Helophorus grandis Illiger, 1798
Hydrophilidae Latreille, 1802
Helochares lividus (Forster, 1771)
Cercyon convexiusculus Stephens, 1829
Cercyon impressus (Sturm, 1807)
Sphaeridium bipustulatum Fabricius, 1781
Sphaeridium lunatum (Fabricius, 1792)
Sphaeridium scarabaeoides (Linnaeus, 1758)
Histeridae Gyllenhal, 1808
Dendrophilus punctatus (Herbst, 1792)
Onthophilus striatus (Forster, 1771)
Margarinotus neglectus (Germar, 1813)
Margarinotus ventralis (Marseul, 1854)
Hister unicolor Linnaeus, 1758
Hydraenidae Mulsant, 1844
Ochthebius minimus (Fabricius, 1792)
Ptiliidae Erichson, 1845
Acrotrichis cognata (Matthews, A., 1877)
Acrotrichis grandicollis (Mannerheim, 1844)
Acrotrichis insularis (Mäklin, 1852)
Acrotrichis rosskotheni Sundt, 1971
Leiodidae Fleming, 1821
Agathidium confusum Brisout de Barneville, 1863
Choleva glauca Britten, 1918
Choleva jeanneli Britten, 1922
Catops fuliginosus Erichson, 1837
Catops nigricans (Spence, 1813)
Silphidae Latreille, 1807
Necrodes littoralis (Linnaeus, 1758)
Silpha tristis Illiger, 1798
Staphylinidae Latreille, 1802
Acidota crenata (Fabricius, 1792)
Anthobium atrocephalum (Gyllenhal, 1827)
Lesteva longoelytrata (Goeze, 1777)
Phyllodrepa floralis (Paykull, 1789)
Metopsia clypeata (Müller, P.W.J., 1821)
Micropeplus staphylinoides (Marsham, 1802)
Bolitobius castaneus (Stephens, 1832)
Coproporus immigrans (Schülke, 2007)
Ischnosoma splendidum (Gravenhorst, 1806)
Lordithon thoracicus (Fabricius, 1777)
Parabolitobius inclinans (Gravenhorst, 1806)
Sepedophilus nigripennis (Stephens, 1832)
Tachinus pallipes (Gravenhorst, 1806)
Tachinus subterraneus (Linnaeus, 1758)
Tachyporus atriceps Stephens, 1832
Tachyporus chrysomelinus (Linnaeus, 1758)
Tachyporus hypnorum (Fabricius, 1775)
Tachyporus nitidulus (Fabricius, 1781)
Tachyporus pallidus Sharp, 1871
Aleochara curtula  (Goeze, 1777)
Aleochara kamila Likovský, 1984
Aleochara sparsa Heer, 1839
Alaobia scapularis  (Sahlberg, C.R., 1831)
Aloconota gregaria (Erichson, 1839)
Atheta aquatica (Thomson, C.G., 1852)
Atheta castanoptera (Mannerheim, 1830)
Atheta harwoodi (Williams, 1930)
Boreophilia eremita (Rye, 1866)
Datomicra dadopora (Thomson, C.G., 1867)
Datomicra nigra (Kraatz, 1856)
Dinaraea angustula (Gyllenhal, 1810)
Plataraea brunnea (Fabricius, 1798)
Encephalus complicans Stephens, 1832
Gyrophaena joyioides Wüsthoff, 1937
Gyrophaena manca Erichson, 1839
Cypha seminulum (Erichson, 1839)
Deleaster dichrous (Gravenhorst, 1802)
Stenus bifoveolatus Gyllenhal, 1827
Stenus brunnipes Stephens, 1833
Stenus clavicornis (Scopoli, 1763)
Stenus flavipes Stephens, 1833
Stenus fulvicornis Stephens, 1833
Stenus ossium Stephens, 1833
Stenus providus Erichson, 1839
Stenus pubescens Stephens, 1833
Stenus solutus Erichson, 1840
Astenus lyonessius (Joy, 1908)
Astenus pulchellus (Heer, 1839)
Lithocharis nigriceps (Kraatz, 1859)
Othius punctulatus (Goeze, 1777)
Philonthus cognatus Stephens, 1832
Philonthus succicola Thomson, C.G., 1860
Philonthus tenuicornis Mulsant & Rey, 1853
Quedius curtipennis Bernhauer, 1908
Quedius maurorufus (Gravenhorst, 1806)
Quedius maurus (Sahlberg, C.R., 1830)
Quedius mesomelinus (Marsham, 1802)
Quedius scintillans (Gravenhorst, 1806)
Ocypus aeneocephalus (De Geer, 1774)
Ocypus fuscatus (Gravenhorst, 1802)
Ontholestes murinus (Linnaeus, 1758)
Ontholestes tessellatus (Geoffroy, 1785)
Platydracus stercorarius (Olivier, 1795)
Tasgius ater (Gravenhorst, 1802)
Tasgius morsitans (Rossi, 1790)
Gauropterus fulgidus (Fabricius, 1787)
Xantholinus longiventris Heer, 1839
Geotrupidae Latreille, 1802
Geotrupes stercorarius (Linnaeus, 1758)
Scarabaeidae Latreille, 1766
Aphodius ater (De Geer, 1774)
Aphodius contaminatus (Herbst, 1783)
Aphodius depressus (Kugelann, 1792)
Aphodius fimetarius (Linnaeus, 1758)
Aphodius foetens (Fabricius, 1787)
Aphodius foetidus (Herbst, 1783)
Aphodius fossor (Linnaeus, 1758)
Aphodius granarius (Linnaeus, 1767)
Aphodius haemorrhoidalis (Linnaeus, 1758)
Aphodius luridus (Fabricius, 1775)
Aphodius pedellus (De Geer, 1774)
Aphodius rufus (Moll, 1782)
Aphodius sticticus (Panzer, 1798)
Hoplia philanthus Welsh Chafer (Füessly, 1775)
Melolontha melolontha Common Cockchafer (Linnaeus, 1758)
Amphimallon solstitale Summer Chafer (Linnaeus, 1758)
Scirtidae Fleming, 1821
Cyphon pubescens  (Fabricius, 1792)
Buprestidae Leach, 1815
Agrilus sinuatus (Olivier, 1790)
Byrrhidae Latreille, 1804
Simplocaria semistriata (Fabricius, 1794)
Byrrhus fasciatus (Forster, 1770)
Byrrhus pilula (Linnaeus, 1758)
Elateridae Leach, 1815
Athous subfuscus (Müller, O.F., 1764)
Agriotes acuminatus (Stephens, 1830)
Sericus brunneus (Linnaeus, 1758)
Cantharidae Imhoff, 1856 (1815)
Cantharis dicipiens Baudi, 1871
Cantharis lateralis Linnaeus, 1758
Cantharis nigra (De Geer, 1774)
Rhagonycha limbata Thomson, C.G., 1864
Rhagonycha testacea (Linnaeus, 1758)
Ptinidae Latreille, 1802
Ochina ptinoides (Marsham, 1802)
Ernobius mollis (Linnaeus, 1758)
Anobium inexspectatum Lohse, 1954
Anobium punctatum (De Geer, 1774)
Lymexylidae Fleming, 1821
Lymexylon navale (Linnaeus, 1758)
Kateretidae Kirby, 1837
Brachypterus glaber (Stephens, 1832)
Nitidulidae Latreille, 1802
Meligethes atratus (Olivier, 1790)
Meligethes flavimanus Stephens, 1830
Omosita discoidea (Fabricius, 1775)
Pocadius adustus Reitter, 1888
Monotomidae Laporte, 1840
Rhizophagus fenestralis (Linnaeus, 1758)
Monotoma bicolor (Villa & Villa, 1835)
Silvanidae Kirby, 1837
Uleiota planata (Linnaeus, 1761)
Silvanus bidentatus (Fabricius, 1792)
Laemophloeidae Ganglbauer, 1899
Cryptolestes duplicatus (Waltl, 1839)
Phalacridae Leach, 1815
Olibrus aeneus (Fabricius, 1792)
Cryptophagidae Kirby, 1837
Telmatophilus caricis (Olivier, 1790)
Cryptophagus denticulatus Heer, 1841
Atomaria lewisi Reitter, 1877
Atomaria linearis Pygmy Beetle Stephens, 1830
Ephistemus globulus (Paykull, 1798)
Erotylidae Latreille, 1802
Triplax aenea (Schaller, 1783)
Byturidae Gistel, 1848
Byturus tomentosus Raspberry Beetle (De Geer, 1774)
Coccinellidae Latreille, 1807
Coccidula rufa (Herbst, 1783)
Scymnus femoralis (Gyllenhal, 1827)
Scymnus frontalis (Fabricius, 1787)
Scymnus schmidti Fürsch, 1958
Nephus redtenbacheri (Mulsant, 1846)
Anisosticta novemdecimpunctata Water Ladybird (Linnaeus, 1758)
Tytthaspis sedecimpunctata 16-spot Ladybird (Linnaeus, 1761)
Myzia oblongoguttata Striped Ladybird (Linnaeus, 1758)
Halyzia sedecimguttata Orange Ladybird (Linnaeus, 1758)
Psyllobora vigintiduopunctata 22-spot Ladybird (Linnaeus, 1758)
Adalia decempunctata 10-spot Ladybird (Linnaeus, 1758)
Harmonia axyridis Harlequin Ladybird (Pallas, 1773)
Subcoccinella vigintiquattuorpunctata 24-spot Ladybird (Linnaeus, 1758)
Corylophidae LeConte, 1852
Orthoperus nigrescens Stephens, 1829
Latridiidae Erichson, 1842
Enicmus histrio Joy & Tomlin, 1910
Corticaria ferruginea Marsham, 1802
Mycetophagidae Leach, 1815
Mycetophagus quadriguttatus Müller, P.W.J., 1821
Typhaea stercorea Hairy Fungus Beetle (Linnaeus, 1758)
Melandryidae Leach, 1815
Orchesia minor Walker, 1836
Melandrya caraboides (Linnaeus, 1761)
Colydiidae Billberg, 1820
Cicones variegatus (Hellwig, 1792)
Tenebrionidae Latreille, 1802
Lagria hirta (Linnaeus, 1758)
Isomira murina (Linnaeus, 1758)
Oedemeridae Latreille, 1810
Oedemera lurida (Marsham, 1802)
Oedemera nobilis (Scopoli, 1763)
Meloidae Gyllenhal, 1810
Meloe proscarabaeus Black Oil Beetle Linnaeus, 1758
Salpingidae Leach, 1815
Rabocerus gabrieli Gerhardt, 1901
Anthicidae Latreille, 1819
Stricticollis tobias (Marseul, 1879)
Scraptiidae Gistel, 1848
Anaspis costai Emery, 1876
Cerambycidae Latreille, 1802
Rhagium mordax (De Geer, 1775)
Stenocorus meridianus (Linnaeus, 1758)
Arhophalus rusticus (Linnaeus, 1758)
Poecilium alni (Linnaeus, 1767)
Chrysomelidae Latreille, 1802
Oulema rufocyanea (Suffrian, 1847)
Cryptocephalus fulvus (Goeze, 1777)
Chrysolina oricalcia (Müller, O.F., 1776)
Chrysolina polita (Linnaeus, 1758)
Chrysolina staphylaea (Linnaeus, 1758)
Phratora laticollis Suffrian, 1851
Galeruca tanaceti (Linnaeus, 1758)
Agelastica alni  (Linnaeus, 1758)
Phyllotreta nemorum (Linnaeus, 1758)
Phyllotreta nigripes (Fabricius, 1775)
Longitarsus luridus (Scopoli, 1763)
Longitarsus melanocephalus (De Geer, 1775)
Altica lythri Aubé, 1843
Altica oleracea (Linnaeus, 1758)
Crepidodera aurea (Fourcroy, 1785)
Crepidodera aurata (Marsham, 1802)
Crepidodera fulvicornis (Fabricius, 1792)
Sphaeroderma rubidum (Graëlls, 1858)
Sphaeroderma testaceum (Fabricius, 1775)
Psylloides chrysocephala (Linnaeus, 1758)
Cassida flaveola Thunberg, 1794
Cassida rubiginosa Müller, O.F., 1776
Anthribidae Billberg, 1820
Platyrhinus resinosus (Scopoli, 1763)
Platystomos albinus (Linnaeus, 1758)
Apionidae Schönherr, 1823
Ceratapion gibbirostre (Gyllenhal, 1813)
Exapion fuscirostre (Fabricius, 1775)
Eutrichapion vorax (Herbst, 1797)
Protapion apricans (Herbst, 1797)
Curculionidae Latreille, 1802
Archarius salicivorus Willow Gall Weevil (Paykull, 1792)
Curculio glandum Acorn Weevil Marsham, 1802
Curculio rubidus (Gyllenhal, 1836)
Anthonomus rubi Strawberry Blossom Weevil (Herbst, 1795)
Cionus alauda (Herbst, 1784)
Cionus hortulanus (Fourcroy, 1785)
Cionus scrophulariae (Linnaeus, 1758)
Dorytomus taeniatus (Fabricius, 1781)
Mecinus pyraster (Herbst, 1795)
Miarus campanulae (Linnaeus, 1767)
Orchestes pilosus (Fabricius, 1781)
Rhamphus oxyacanthae (Marsham, 1802)
Ceutorhynchus constrictus (Marsham, 1802)
Ceutorhynchus pallidactylus Cabbage Stem Weevil (Marsham, 1802)
Ceutorhynchus picitarsis Gyllenhal, 1837
Thamiocolus viduatus Thomson, C.G., 1859
Euophryum confine (Broun, 1881)
Acalles misellus Boheman, 1844
Acalles ptinoides (Marsham, 1802)
Kyklioacalles roboris (Curtis, 1835)
Phyllobius pomaceus Green Nettle Weevil Gyllenhal, 1834
Phyllobius roboretanus Small Green Nettle Weevil Gredler, 1882
Phyllobius virideaeris (Laicharting, 1781)
Liophloeus tesselatus (Müller, O.F., 1776)
Sciaphilus asperatus (Bonsdorff, 1785)
Sitona lepidus Gyllenhal, 1834
Sitona striatellus Gyllenhal, 1834
Romualdius angustisetulus Hansen, 1915
Hylobius abietis Pine Weevil (Linnaeus, 1758)
Pissodes pini (Linnaeus, 1758)
Pityophthorus pubescens (Marsham, 1802)
Kissophagus hederae (Schmitt, 1843)
Hylurgops palliatus (Gyllenhal, 1813)