
| Cryptocephalus coryli | ||
| At Sherwood Forest NNR 2008-2011 | ||
| In 2008, we accidentally
rediscovered the RDB1 and UKBAP priority beetle Cryptocephalus
coryli at a single location within the Sherwood
Forest NNR, which is currently believed to be the only
site in the UK where this beetle still occurs. Since 2008, we have continuously surveyed for the beetle from April to July each year, in order to monitor the population at Sherwood, producing short summaries of the records and detailing any findings. These can be found by scrolling further down this page, where there is a complete list of all known records from the Sherwood Forest NNR. 2011 turned out to be an exceptional year for numbers of adult Cryptocephalus coryli (more commonly known as the Hazel Pot Beetle) and by employing two unique survey techniques, we made several important discoveries regarding the habits of this endangered beetle. |
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| Life
history of Cryptocephalus coryli
and other Cryptocephalus
species The life cycle of Cryptocephalus beetles is an interesting one. Once mated, the female begins to lay, holding each egg laid between her rear tarsi, turning it and covering it with dung at the same time. Once the egg is completely coated, the female drops the 'pot' to the ground. When the egg hatches, the young larva makes a hole at one end, through which it can emerge to feed and move about in the leaf litter. As the larva grows, it enlarges the pot which eventually attains a length of about 14mm. In the wild, larval development usually takes two years, but larvae are often full grown within a year when kept in captivity. Once mature, the larva seals the end of the pot, turns around inside to face the other way and pupates. When the beetle emerges, it cuts itself out of the pot and climbs the nearest vegetation before flying. |
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| Historical
records of Cryptocephalus coryli
in Nottinghamshire and the UK Many years ago, Cryptocephalus coryli was a widespread species across the southern counties of England, but it declined greatly during the last century to become one of the UK's rarest insects. It's decline is thought to have coincided with the reduction in coppicing. Since 1970 it had been found at only three single sites - in Berkshire, Lincolnshire and Surrey (Ref: Natural England). In the south of the UK, it occurred on hazel growing along woodland edges, rides and hedgerows, but in the north preferred a heathland habitat with Birch scrub or young trees. Historically, Sherwood Forest was always regarded as being good for C. coryli by collectors in the early 1900's, when over a dozen adults could be taken on each visit to this locality, with the beetles either being beaten or picked from young Birch and Oak during June. |
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Some Sherwood Forest
records dating back to the 1930's, remained something of
a mystery until early in 2011, when work to re-organise
the coleoptera collection at Leicestershire Museum CRC at
Barrow-upon Soar, finally helped clear up some of the
confusion regarding records dating back to around this
time. The re-organisation also provided additional records dating from 1929, 1939 and 1940, which seemed to have been previously unknown or were lost from the record books. These are listed in the table of all Sherwood's C. coryli records at the bottom of this page. Although most of these records still list no recorder or determiner, it is hoped that we can make further enquiries to complete the relevant information over the next few months. In May 28th 2008, we found an attractive red beetle resting on a dead Rosebay Willowherb stem in an area of the Sherwood Forest Country Park. It's identity as a female Cryptocephalus coryli, was determined from photographs took on site, on our return home. |
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| Being amazed
at the potential of the find and after reading about the
reintroduction scheme that had taken place in
Lincolnshire in 2000, we made further enquiries to
ascertain whether a similar introduction had occurred in
Nottinghamshire, and in particular at Sherwood Forest. It
turned out that no such introduction scheme had taken
place and that this record was the first in
Nottinghamshire for at least 70 years. Two more records
of females followed on May 29th and June 4th. The area was surveyed on behalf of Natural England East Midlands by Allan and Annette Binding in 2009, whilst we continued our own searches for the beetle. Both the Bindings and ourselves, also conducted unproductive surveys at other potentially suitable locations within the Country Park area. We surveyed the area again in 2010 on behalf of Nottinghamshire County Council and in conjuction with our own survey work, finding only two adults, despite what seemed very favourable weather conditions. |
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| In Autumn 2009, we were
consulted about the potential affects thinning work (as
part of the sites on-going management plan) in the two
main compartments bordering and containing C. coryli
would have. In the end it was decided that work
in one compartment would be postponed to a later date,
with work being carried out in the other during early
March 2010. Thinning work and the removal of timber was carried out as per our recommendations and damage to habitat by machinery was minimal at most. Birch and Hawthorn scrub known to be used by C. coryli females was left untouched. Trees in the remaining compartment not touched in March 2010, were later thinned in January 2011. We were personally involved with the work, selecting and marking the trees to be felled. The whole operation went very well, with an increase in potentially suitable areas of habitat being created and opened up, but with just two adults recorded in 2010, it was always difficult to know if the beetle was in decline or it had just been a bad year. In the end, 2011 turned out to be a remarkable year with a total of 71 adults recorded. |
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| Some
new observations on the habits of Cryptocephalus
coryli during 2011 Continued study and survey of the site during 2011 saw us greatly increase our personal understanding of the habits of C. coryli and several important new discoveries were made, which would certainly go some way towards explaining why coryli went 'missing' at Sherwood Forest (and probably other sites in the UK) for decades without being found. Two adults observed on small scrub Oak and Birch and then seen flying directly to the top of nearby mature Birches, provided us with our first indication that we may have spent many hours looking in the wrong place (although still proving successful) and that maybe we were not looking high enough. We were able to confirm this during May and early June by advancing our survey techniques, conducting several surveys of the larger trees from the ground using binoculars and a telescope, all of which produced double figure counts during peak season. It was a slow and painstaking method involving several scans of each tree, but proved extremely successful on windless days. Finding that adults generally preferred taller trees, could be a likely expanation for the poor numbers recorded in 2010. |
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It was found that
branches of thicker foliage, with a sunny aspect and
around 30 feet above ground level were particularly
favoured. No adults were found at the very top of the
largest Birches in the area we surveyed using a cherry
picker to a height of 50 feet, but it was obvious that
the foliage quality on larger Birches was of noticeably
poor quality. During windy weather, adult coryli adopt a policy of moving from the leaves and wrapping their legs around thin stems until conditions have settled. Even when on small scrub, adults are actually quite difficult to find when doing this. We were hampered by very breezy conditions on the day of the survey, but still recorded 13 adults which would not have been visible from the ground surveying with binoculars and telescope. Over the years, the available literature has suggested that C. coryli uses 'key' trees, typically Birch scrub and young trees and whilst females are often found on Birches fitting that criteria, the number of adults found on these must be a fraction of the actual population. |
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| We now think
that perhaps too much emphasis has been placed on the use
of 'key' trees in the past. But if these trees are
neccessary for coryli, then 'key' groups of
mature Birches are more likely, where the microclimatic
conditions for the beetle are optimal and the foliage is
healthy. Evidence for this comes from recording loose
groups of adults within close proximity on single or
adjoining trees, though we have recorded three adults on
a single leaf. Our reasoning for this stems from the fact that key trees have always been habitually small and much easier to check for coryli than much larger (30-50 feet high) trees. We believe that this has led to the misconception that key trees are used by successive generations of females. We are inclined to think that after spending many hours checking several key trees at Sherwood CP, we would have expected to have noted males flying around them or sat on them in their search for females. Several 'key' trees used in 2008 and 2009 have not been used since and it is most likely that many adults found on small scrub, have just simply climbed there after emergence, as they do grass and other vegetation before their first flight. |
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| Although
males are more active than females near the tops of
Birches, we noted that they spend much time sunning
themselves on warm afternoons. Greatest male flight
activity seems to occur between 09:00h and 12:00h on fine
warm days. We have never recorded males on scrub Birches
and have found no evidence yet to suggest that females
descend from height deliberately seeking out scrub (key?)
Birch. We also recorded a female on Downy Birch (Betula
pubescens) for the first time in 2011. Plotting the locations of all adult coryli found at Sherwood Forest since 2008 on the map above, certainly shows that although this beetle is fussy about its habitat requirements, having the correct habitat for larval devlopment is even more important. The lack of habitat which combines the two required elements for both adult and larval stages, helps explain why coryli remains restricted at Sherwood Forest CP. Slightly damper soil containing a mixture of coarser grasses, Rosebay Willowherb and (to a lesser extent) Umbellifers, seems to be a key factor in larval development. Areas of finer grasses such as Fescues on drier soils are less suitable unless the grass is left untouched. |
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| Summary of Cryptocephalus coryli records from Sherwood Forest NNR in 2011 | ||
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| Following on from a poor
year for C. coryli numbers, coupled with an
exceptionally cold period at the end of 2010, it was
always going to be a guess as to what would happen in
2011. Certainly the weather during the early months of
this year was especially favourable, with the remainder
of the Winter and early Spring being very dry. April's
dry and at times very warm weather, saw many
invertebrates produce early first dates and the first
adult coryli (a male) was found on the
exceptionally early date of April 23rd. This was
certainly the earliest record for Sherwood Forest and
very early nationally, with mid-May to early June
generally being regarded as the main emergence period for
adults. It was May 1st before the first female was found on a small Oak, but this was still another early date. She was photographed, but very quickly took flight and flew high up to the top of a nearby mature Birch. We made a return visit to the site in the late afternoon, to mark the locations of this year's two C. coryli, and found a third adult (a male) on a lower branch of a large Birch. |
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| May 2nd saw a further two females found on a small Oak (which soon flew off unseen) and a large Birch respectively and within a few feet of each other. Scanning the lower branches of large Birches growing along the southern edge of the favoured area with binoculars on May 3rd, revealed another male and two females, bringing the total of adults for the year to eight. | ||
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May 4th was a significant
day, when a concerted effort was made to check the higher
branches of the mature Birches along the edge of the
favoured plantation. Binoculars and a Kowa telescope were
used to find and sex any adults, resulting in a
staggering total of 11 adults including six females and
five males. One Birch alone produced three males in close
proximity to each other. Perhaps a more important
discovery, was a male and female found in a new location,
slightly away from the known population. This new
location provided additional records of females found on
May 6th and then recently hatched on May 10th. A technique employed by adults in windy conditions, came to light on May 8th. After scanning the upper reaches of Birches with binoculars and a scope revealed no adults, the reason for this was made clear by this female (shown left) found on a small scrub Birch, who was literally clinging on to the stem rather than the usual method of sitting directly on the leaf in fine conditions. Even with their red colouration, C. coryli are surprisingly difficult to see when adopting this position on small scrub Birch and Oak. By May 8th, the total number of adults found had reached 26, with the percentage of females being greater. |
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On May 9th we were able
to conduct the first of two unique surveys with the very
kind help and support of WKW Tree Services. The surveys
were based at tree top height, using a cherry picker
operated for us by the extremely willing Matt Vaughan. At the end of six hours, a total of 14 C.coryli had been located, which was a new record total, despite the windy conditions. All but one were at the very tops of mature Birches lining the southern edge of the favoured location, including one mating pair. With the wind being so gusty at times, all of the beetles were slightly down the Birch stems, clinging on tightly in the manner we noted the previous day when conditions were similar. A more thorough detailing of the days results can be found via the links at the bottom of this page. The period May 11-17th continued to provide more records, with the total adults for the year eventually reaching 50 on the latter date. This period also brought with it slightly unsettled weather, cooler and frequently too windy to check the tops of Birches for adults, but also provided a string of records of recently hatched adults on either Willowherb or grass. |
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| It seemed
obvious that newly emerged adults will hatch in slightly
cooler conditions, but remain on the grass etc, until
conditions are warm enough for flight. Probably the most
pleasing of all our coryli finds during this
period, was eventually finding a half-grown larva and pot
in leaf litter on May 16th, directly underneath a newly
emerged male. This was a personal triumph for us after
spending many fruitless hours searching leaf litter and
grass tussocks during the late Winter and early Spring
period. During the second half of May, conditions (although largely dry and sunny) were often windy, so it was not possible to survey the higher Birch foliage until early June. A few adults still kept turning up though, mostly on one clump of scrub Birch, but we also several found newly hatched adults on grass. On June 2nd, the weather was finally still, sunny and warm, so a couple of hours were spent scanning the tops of Birches with binoculars and a scope. A total of 15 adults were located (eight males and seven females) mostly very high up. Notable were two males and a female on the same leaf at the top of a Birch at the eastern end of the Gleadthorpe open, a grassy area within the country park. There seemed to be a sudden end to the adult season, with just one solitary female with a damaged elytra, found in another new area on June 9th and remaining on the same scrub Birches until June 24th. Largely windy conditions continued, making scanning with binoculars and a telescope impossible, but also proving to be surprisingly fruitless when it was calm enough. A second tree top survey using the cherry picker was again conducted at the end of July and whilst we knew that finding any coryli was always doubtful, it was still worth the effort to check. A total of 71 adults were found during 2011. |
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| Summary of Cryptocephalus coryli records from Sherwood Forest NNR in 2010 | ||
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| Despite having the
coldest Winter for many years, we began actively
searching for the first C. coryli from around
the start of the last week of April 2010. The weather hampered searches for any very early adults and despite some fine sunny days there were overnight temperatures down to -3 during early May. As in 2009, all scrub Oaks were affected by frost, meaning that at least if any beetles were to emerge, then they would most likely be on grass, or on the limited amount of scrub Birch and Hawthorn (both unaffected by frost and in full leaf) available on site. On May 16th, some three weeks after the start of our search, we found the first female on a small scrub Birch along a south-facing section of Birch woodland. This remained the only record until June 19th, when a male was found on young Oak growth on a stump cut down during the Winter. |
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| Summary
of Cryptocephalus coryli
records from Sherwood Forest NNR in 2009 The first adults (four) were found on May 10th, but casual searching actually commenced at the end of April. Hatching of the adults seemed to coincide with fine days, on which the temperature was fairly warm. Many adults were found on grass, including one female which was photographed egg-laying whilst on grass, both observations which had not been recorded before. Between May 10th and June 12th, we recorded a total of 13 adults of which four were males. These are listed at the bottom of this page. Further adults were recorded during a targeted survey by Allan and Annette Binding. |
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Adults were found over a
larger area than in 2008, but still in a restricted
section of Sherwood Forest CP. Records throughout the
Spring indicated that this beetle is largely found along
the southern and eastern edges of one section of Birch
and Oak woodland within Sherwood Forest CP, with adults
generally being noted on grass (providing most records)
or on Hawthorn and Birch scrub. The number of adults found during 2009 indicates that Sherwood has a very healthy population of C. coryli and an appeal for more information by Allan and Annette Binding on the few other known UK sites for this beetle, saw no response. It appears that known sites in Lincolnshire have also proved negative for C. coryli. Below:- Two C. coryli females, photographed at Sherwood Forest CP on June 8th 2009. The darker one of the two was actually found on June 4th and retained in captivity during a period of very wet weather. On release, we were fortunate to find this second female on a small Birch nearby. |
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| Summary
of Cryptocephalus coryli
records from Sherwood Forest NNR in 2008 There were a total of three records of C. coryli from Sherwood Forest in 2008. All three related to females, one of which spent several days on a small Birch and was photographed egg-laying on at least one occasion. The records and notes on their discovery are listed below. |
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| Following the discovery
and identification confirmation of the first female on
28/05/08, further visual searches of Birch/Oak scrub
foliage in other nearby compartments were made. These
took place on an occasional basis until late June. Visual searches soon proved successful on a further two occasions, with another found a day after the first on 29/05/08 and a third, a few days later on 04/06/08. This female remained present till 07/06/08, never leaving her favoured Birch twig and was photographed in the process of egg-laying on at least one occasion. The favoured Birch was just less than a metre tall, but just over a metre wide. Adjacent scrub was predominantly Oak of various sizes and a single small Hawthorn. Across the path there is an area of open, grassy heathland, in which there are several small areas of Birch scrub. These could be utilised as suitable sites for egg-laying females, but may be situated too far from the woodland edge. |
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| It is possible that the Oak scrub were ignored by the females, due to being severely frost damaged after leaf emergence. By the time the Oaks had recovered, the C. coryli emergence/breeding period was over. The Birch also held numbers of Sloe Bug (Dolycoris baccarum) and the ant (Formica fusca). No prolonged observations were made at the time to determine whether these two insect species had any adverse effects on the female coryli presence, but any effects must have been minimal given the female's lengthy duration of stay. | ||
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| All known Sherwood Forest NNR records of Cryptocephalus coryli |
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| 1903 | .. | Kidson Taylor, J. | .. | N/A | .. | .. | Sherwood Forest | ||||
| 1904 | .. | Kidson Taylor, J. and Hardy. | .. | N/A | .. | .. | Sherwood Forest | ||||
| 1906 | .. | Kidson Taylor, J. | .. | N/A | .. | .. | Sherwood Forest | ||||
| 24/06/1907 | .. | Donisthorpe, H. | .. | N/A | .. | .. | Sherwood Forest | ||||
| 25/06/1907 | .. | Donisthorpe, H. | .. | N/A | .. | .. | Sherwood Forest | ||||
| 1908 | .. | Bedwell, E.C. | .. | N/A | .. | .. | Sherwood Forest | ||||
| 05/06/1929 | .. | No known recorder | .. | N/A | .. | .. | Sherwood Forest. Adult | ||||
| 22/05/1938 | .. | Tozer, D. | .. | N/A | .. | .. | Sherwood Forest. Adult | ||||
| 29/05/1938 | .. | No known recorder | .. | N/A | .. | .. | Sherwood Forest. Adult | ||||
| 05/06/1938 | .. | Tozer, D. | .. | N/A | .. | .. | Sherwood Forest. Adult | ||||
| 12/06/1938 | .. | Tozer, D. | .. | N/A | .. | .. | Sherwood Forest. Adult | ||||
| 26/06/1938 | .. | No known recorder | .. | N/A | .. | .. | Sherwood Forest. Two adults | ||||
| 03/05/1939 | .. | No known recorder | .. | N/A | .. | .. | Sherwood Forest. Adult | ||||
| 17/06/1940 | .. | No known recorder | .. | N/A | .. | .. | Sherwood Forest. Adult | ||||
| 27/06/1940 | .. | No known recorder | .. | N/A | .. | .. | Sherwood Forest. Two adults | ||||
| 28/05/2008 | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On grass | ||||
| 29/05/2008 | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On isolated small Hawthorn. | ||||
| 04/06/2008 | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On Birch scrub adjacent to Birch/Oak woodland. Photographs showed this female to be egglaying. Still present on same Birch over the next few days and occasionally more active. | ||||
| 10/05/2009 | .. | Pendleton, T.A. and D.T. | .. | Male | .. | .. | On grass at 11:30h (presumed recently hatched) | ||||
| ..... | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On small scrub Birch used in 2008. Found at 12:10h, then flew off. | ||||
| ..... | Pendleton, T.A. and D.T. | Female | On small Hawthorn used in 2008 at 14:30h. Remained in situ till 18/05/2009, only moving when the weather warmed up. | ||||||||
| ..... | .. | Pendleton, T.A. and D.T. | .. | Male | .. | .. | On grass at 15:50h (presumed recently hatched) | ||||
| 12/05/2009 | .. | Binding, A.E. and A. | .. | Female | .. | .. | On leaf of mature Birch (12 feet above ground level) | ||||
| ..... | .. | Binding, A.E. and A. | .. | Female | .. | .. | On small Hawthorn used in 2008 | ||||
| 18/05/2009 | .. | Binding, A.E. and A. | .. | Female | .. | .. | On small Hawthorn used in 2008 | ||||
| 20/05/2009 | .. | Pendleton, T.A. and D.T. | .. | Male | .. | .. | On small Oak at 12:30h (presumed recently hatched) | ||||
| 21/05/2009 | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On Hawthorn from late morning till early afternoon, but not seen later. | ||||
| ..... | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On grass at 14:00h. Relocated by hand to same Hawthorn as above female. | ||||
| ..... | .. | Binding, A.E. and A. | .. | Female | .. | .. | On grass at 16:35h | ||||
| 23/05/2009 | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On grass at 14:45h, then flew to nearby small Oak | ||||
| 24/05/2009 | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On grass | ||||
| ..... | .. | Binding, A.E. and A. | .. | Male | .. | .. | Observed in flight | ||||
| 04/06/2009 | .. | Pendleton, T.A. and D.T. | .. | Female | .. | On grass at 15:00h. Proved to have mated and taken into captivity. Released in same locality on June 8th. | |||||
| 08/06/2009 | .. | Pendleton, T.A. and D.T. Binding, A.E. and A. | .. | Female | .. | On small Birch, remained until 11/06/2009 | |||||
| 11/06/2009 | .. | Binding, A.E. and A. | .. | Female | .. | .. | On small Birch | ||||
| 12/06/2009 | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On grass at 11:46h | ||||
| ..... | .. | Pendleton, T.A. and D.T. | .. | Male | .. | .. | On grass at 14:36h | ||||
| 16/05/2010 | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | Found at 11:35h on small scrub Birch | ||||
| 19/06/2010 | .. | Pendleton, T.A. and D.T. | .. | Male | .. | Found at 11:05h resting on young Oak growth from stump cut down the previous Autumn. Later observed feeding on leaves at 12:35h. Weather fairly cool and breezy | |||||
| 23/04/2011 | Pendleton, T.A. and D.T. | Male | On grass at 10:20h. The first ever April record of C. coryli from the Sherwood Forest NNR and possibly the earliest ever national record. Relocated on nearby Hawthorn on 24/04/2011 | ||||||||
| 01/05/2011 | Pendleton, T.A. and D.T. | Female | Found on a small Oak at 12:05h, but soon became active and flew high to the top of a nearby Birch. | ||||||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Male | .. | .. | On lower branch of large Birch, found at 15:40h. Present again on 02/05/2011. | ||||
| 02/05/2011 | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On Oak growing underneath/through a large Birch, found at 13:15h. | ||||
| Pendleton, T.A. and D.T. | Female | Found within a few feet of above female, on lower branch of a large Birch, found at 13:20h. | |||||||||
| 03/05/2011 | .. | Pendleton, T.A. and D.T. and Joynt, G. | .. | Male | .. | .. | 12 feet up large Birch at 15:00h | ||||
| .. | .. | Pendleton, T.A. and D.T. and Joynt, G. | .. | Female | .. | .. | 16 feet up large Birch at 15:00h | ||||
| .. | .. | Pendleton, T.A. and D.T. and Joynt, G. | .. | Female | .. | .. | 16 feet up large Birch at 15:20h | ||||
| 04/05/2011 | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On scrub Birch at 12:42h. | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On small Oak at 12:46h. | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On lower branches (7 feet) of mature Birch at 12:53h | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On lower branches (9 feet) of mature Birch at 13:15h | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Male | .. | .. | On branches (30 feet) of mature Birch at 14:00h | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Male | .. | .. | On branches (30 feet) of mature Birch at 14:00h | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Male | .. | .. | On branches (30 feet) of mature Birch at 14:00h | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Male | .. | .. | On lower branches (10 feet) of mature Birch at 14:15h | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On branches (30 feet) of mature Birch at 14:25h | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Male | .. | .. | On lower branches (12 feet) of mature Birch at 14:45h | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On branches (22 feet) of mature Birch at 15:00h | ||||
| 05/05/2011 | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On lower branches (7 feet) of mature Birch at 16:35h | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On lower branches (6 feet) of mature Birch at 16:38h, on same tree as above female | ||||
| 06/05/2011 | .. | Pendleton, D.T. and Joynt, G. | .. | Male | .. | .. | On branches (30 feet) of mature Birch at 14:45h, observed mating with female | ||||
| .. | .. | Pendleton, D.T. and Joynt, G. | .. | Female | .. | .. | On branches (30 feet) of mature Birch at 14:45h, observed mating with above male | ||||
| .. | .. | Pendleton, D.T. and Joynt, G. | .. | Female | .. | .. | On branches (20 feet) of mature Birch at 15:15h | ||||
| 07/05/2011 | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On branches (35 feet) of mature Birch at 13:30h | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On scrub Oak at 13:40h | ||||
| 08/05/2011 | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On scrub Birch at 12:27h. Still present on 12/05/2011. | ||||
| 09/05/2011 | .. | Pendleton, T.A. and D.T. and Vaughan, M. | .. | Female | .... | .. | On scrub Birch. | ||||
| .. | .. | Pendleton, T.A. and D.T. and Vaughan, M. | .. | Female | .. | .. | On branches (30 feet) of mature Birch | ||||
| .. | .... | Pendleton, T.A. and D.T. and Vaughan, M. | .. | Female | .. | .. | On branches (30 feet) of mature Birch | ||||
| .. | .. | Pendleton, T.A. and D.T. and Vaughan, M. | .. | Male | .. | .. | On branches (30 feet) of mature Birch | ||||
| .. | .. | Pendleton, T.A. and D.T. and Vaughan, M. | .. | Female | .. | .. | On branches (35 feet) of mature Birch, observed mating with male | ||||
| .. | .. | Pendleton, T.A. and D.T. and Vaughan, M. | .. | Male | .. | .. | On branches (35 feet) of mature Birch, observed mating with female | ||||
| .. | .. | Pendleton, T.A. and D.T. and Vaughan, M. | .. | Male | .. | .. | On branches (35 feet) of mature Birch | ||||
| .. | .. | Pendleton, T.A. and D.T. and Vaughan, M. | .. | Male | .. | .. | On branches (20 feet) of mature Birch | ||||
| .. | .... | Pendleton, T.A. and D.T. and Vaughan, M. | .. | Male | .... | .. | On branches (24 feet) of mature Birch | ||||
| .. | .. | Pendleton, T.A. and D.T. and Vaughan, M. | .. | Female | .. | .. | On branches (30 feet) of mature Birch | ||||
| .... | .. | Pendleton, T.A. and D.T. and Vaughan, M. | .. | Female | .. | .. | On branches (30 feet) of mature Birch | ||||
| .. | .. | Pendleton, T.A. and D.T. and Vaughan, M. | .. | Female | .. | .. | On branches (20 feet) of mature Birch | ||||
| .. | .. | Pendleton, T.A. and D.T. and Vaughan, M. | .. | Male | .. | .. | On branches (22 feet) of mature Birch | ||||
| 10/05/2011 | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On grass at 12:45h. | ||||
| 11/05/2011 | .. | Pendleton, T.A. and D.T. | .. | Male | .. | .. | On branches (30 feet) of mature Birch, then flew off | ||||
| 12/05/2011 | .. | Pendleton, T.A. and D.T. | .. | Male | .. | .. | On branches (25 feet) of mature Birch | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On branches (15 feet) of mature Birch | ||||
| 13/05/2011 | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On scrub Birch. Still present on 15/05/2011 | ||||
| 15/05/2011 | .. | Pendleton, T.A. and D.T. | .. | Male | .. | .. | On Rosebay Willowherb stem within grass in overcast weather | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On dead Rosebay Willowherb stem within grass in overcast weather | ||||
| 16/05/2011 | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On grass in overcast weather, moved to scrub Oak. Killed by a spider on 19/05/2011. | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Larva | .. | .. | In leaf litter at location of above female | ||||
| 17/05/2011 | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On scrub Birch. Still present on 18/05/2011. | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Male | .. | .. | On grass in overcast weather. Still present on 18/05/2011. | ||||
| 18/05/2011 | .. | Pendleton, T.A. and D.T. | .. | Larva | .. | .. | In leaf litter within small excavated trench | ||||
| 19/05/2011 | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On scrub Downy Birch, then flew off unseen | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Male | .. | .. | On grass, became active and flew high to the top of a mature Birch | ||||
| 25/05/2011 | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On scrub Birch. Noted egg-laying. | ||||
| 31/05/2011 | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On scrub Birch. | ||||
| 31/05/2011 | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On scrub Downy Birch. | ||||
| 02/06/2011 | .. | Pendleton, T.A. and D.T. | .. | Male | .. | .. | On branches (40 feet) of mature Birch | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On branches (30 feet) of mature Birch | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On branches (40 feet) of mature Birch | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Male | .. | .. | On branches (40 feet) of mature Birch | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Male | .. | .. | On branches (35 feet) of mature Birch | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Male | .. | .. | On branches (40 feet) of mature Birch | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On branches (28 feet) of mature Birch | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Male | .. | .. | On branches (28 feet) of mature Birch | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Male | .. | .. | On branches (30 feet) of mature Birch | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Male | .. | .. | On branches (35 feet) of mature Birch | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On branches (35 feet) of mature Birch | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Male | .. | .. | On branches (40 feet) of mature Birch. On same leaf as another male and a female. | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Male | .. | .. | On branches (40 feet) of mature Birch. On same leaf as another male and mating with female. | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On branches (40 feet) of mature Birch. On same leaf as above two males, mating. with one male. | ||||
| .. | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On branches (30 feet) of mature Birch | ||||
| 09/06/2011 | .. | Pendleton, T.A. and D.T. | .. | Female | .. | .. | On scrub Birch. This female had slight damage to the elytra and was present on same scrub Birch on 24/06/2011. | ||||